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Dreaming … dreaming to someday soar free, knowing full well, that the free sky is forever barred to them. In a distant future, after Earth has been devastated and most civilisation ceased to exist, the remaining humans built a moving fortress city called “Plantation” in order to survive and keep some civilisation intact. The children of the “Plantation” live in a pilot training facility best known as the “bird cage.” Not knowing the outside world or the free sky, they are trained for only one purpose: to fight the mysterious enemy that reduced humanity to this miserable state. Learning to pilot what is known as “Franxx” and being trained to fight an enemy they have never even seen, the children believe that only piloting the “Franxx” can prove their existence. Hiro—code number 016—was once hailed a child prodigy, but when he fell behind and wasn’t able to ride a “Franxx,” his existence had lost all meaning. Atfer all … if you can’t pilot a “Franxx,” you might as well not exist at all. One day, however, a mysterious girl called Zero Two appears before him with two lustrous horns rising from her head. A familiar smile flickering across her lips, as she calls out to him: “I found you, my darling.”

The story takes place in Akihabara, Tokyo. Rintarou Okabe, a self-proclaimed mad scientiest, is an eccentric person who heads a Futuristic Gadget Research Laboratory. Together with his friends, Mayuri Shiina and Itaru Hashida, he invents all sorts of things that, in actual fact, often don't have a practical use. Later, he accidently builds a time machine, the Phone Microwave. Contrary to his very own belief, the time machine actually works: From now onwards, he is able to send short mails through time to change the past and, eventually, affect the present. Over the course of the story, Okabe Rintarou admits many other people as new members of his Future Gadget Lab like Kurisu Makise, an extremely intelligent 18-year-old scientist. Soon afterwards, the organisation SERN, that also does research on time travel, becomes aware of Rintarou and his effort to revolutionize the process of travelling through time.